Darcy Graham scored two tries, one either side of half-time, for the Scottish outfit, while Blair Kinghorn – who added three conversions - and Mark Bennett also dotted down.

Elsewhere, Benetton Rugby triumphed over South African side Stormers 22-18 at the Stadio Comunale di Monigo.

Tries from Gianmarco Lucchesi, Ivan Nemer and Tommaso Menoncello helped the Italians recover from a 18-7 deficit midway through the second half.

In the Gallagher Premiership, five-time European Rugby Challenge Cup semi-finalists Newcastle Falcons earned a hard-fought 20-13 victory at Bath Rugby thanks to first-half tries from Adam Radwan, Iwan Stephens and George McGuigan.

Worcester Warriors took defending champions Harlequins to the limit in a 35-29 defeat - Ollie Lawrence, Willi Heinz, Will Chudley and Marc Thomas all scoring tries.

In the TOP 14, Section Paloise edged Montpellier Hérault Rugby 23-22 thanks to tries from Youri Delhommel and Aminiasi Tuimaba in a dramatic final ten minutes.

USAP defeated RC Toulon 12-9 in a kick-fest at the Stade Aimé Giral. Perpignan’s Melvyn Jaminet outbooted Toulon’s Louis Carbonel by four penalties to three to seal victory.

Lyon put up a strong fight against Racing 92 before succumbing 24-20, late tries from Temo Mayanavanua and Baptiste Couilloud proving insufficient.

CA Brive were in their contest against Union Bordeaux-Bègles until half-time, when they led 10-9, but eventually crashed to a 29-10 defeat.

Biarritz Olympique suffered a 59-17 drubbing at the hands of Stade Rochelais, replacement Vincent Martin scoring a try for the Challenge Cup contenders in addition to a penalty try.

2015 European Rugby Challenge Cup runners-up Edinburgh Rugby opened their United Rugby Championship campaign with a 26-22 home victory over Scarlets on Saturday.