The Saracens prop, who won the trophy with the English club in 2016, 2017 and 2019, is one of three changes made by head coach Warren Gatland to the side who won the first Test last weekend after an impressive second-half display as a replacement.

He displaces Worcester Warriors prop Rory Sutherland, while the other changes see Munster Rugby scrum-half Conor Murray take over from Glasgow Warriors’ Ali Price and Gloucester Rugby centre Chris Harris oust Saracens’ Elliot Daly for his first Lions Test cap.

Sutherland, Price and Daly all drop to the bench, where they are joined by Bath Rugby No.8 Taulupe Faletau, who is added to the replacements from last week.

“As always, selection was incredibly tough,” said Gatland. “However, we’ve made the changes we think are the right calls for the weekend’s game.

“It’ll be another tight contest. We know the Springboks will be hurting and they’ll throw everything at us on Saturday, but I think there’s plenty more to come from us too.

“We feel we can go up another level from where we were in the first Test and I would expect us to improve.

“It’s the biggest game on the Tour and we have to embrace the expectation that comes with it. As players and coaches, these are the games you want to be involved in. We’re relishing the opportunity to face the Springboks again on Saturday and potentially seal a series win.”

Springboks head coach Jacques Nienaber has handed Leicester Tigers No.8 Jasper Wiese a first Test start as he comes in to replace Kwagga Smith. Wiese scored three tries in Leicester's run to the Challenge Cup final last season, including against Ulster Rugby and Montpellier Hérault Rugby in the semi-final and final respectively.

Triple European Cup winner Mako Vunipola has been promoted to the British and Irish Lions starting XV for their second Test against South Africa in Cape Town on Saturday.