Lloyd notched a try, a penalty and seven conversions for a 22-point haul on his first start for Bristol Bears as they saw off Zebre Rugby 59-21 at Ashton Gate in Pool 4 - racking up the biggest points total of the weekend. Brive are a point behind them after first-half tries from Vasil Lobzhanidze and Otari Giorgadze laid the platform for a 27-11 win at Stade Francais Paris.

Basham scored a first-half hat-trick for Dragons in their 31-17 home win over Castres Olympique, with Dafydd Howells also crossing before half-time to give the Welsh side a bonus point before the game was half over. Elsewhere in Pool 1, Worcester Warriors also achieved a five-point maximum as Jamie Shillcock claimed three of their nine tries in a 57-14 victory against Enisei-STM in Krasnodar.

History was made in Pool 5 when Leicester Tigers played their first ever match in the Challenge Cup, with winger Holmes grabbing four tries and several England players returning from World Cup duty in their 41-20 home win over Pau. In the same pool, two-time winners Cardiff Blues also got a bonus-point win (38-16) at Rugby Calvisano.

In Pool 2, RC Toulon are sole leaders after South African prop Marcel van der Merwe registered a rare brace of tries to help the French club earn a 20-13 bonus-point win at Bayonne. Toulon are just ahead of Scarlets, who held on for a tense 20-16 home victory over London Irish.

In Pool 3, Edinburgh Rugby and Bordeaux-Bègles both recorded bonus-point victories. Edinburgh chalked up a 31-10 success at Agen, where hooker Cameron Fenton scored a last-gasp fourth try for the Scottish side, and Bordeaux beat Wasps 40-30 at home, with Romain Buros' try two minutes from the end clinching the win.

Ioan Lloyd, Taine Basham and Jonah Holmes were among the star individual performers in Round 1 of the European Rugby Challenge Cup, which produced some historic moments and plenty of drama.